Wales head coach Warren Gatland has made 13 changes to the starting line-up for Saturday’s Rugby World Cup Pool C encounter against Portugal in Nice.

Gatland retains just two of the side – wing Louis Rees-Zammit and number eight Taulupe Faletau – that overcame Fiji 32-26 in a ferocious contest last weekend.

Co-captain Dewi Lake leads Wales after recovering from a knee injury, while scrum-half Tomos Williams wins his 50th cap and there are first World Cup starts for the likes of centre Mason Grady and lock Christ Tshiunza.

Lake suffered a knee problem during Wales’ World Cup warm-up game against England, but he now returns to pack down alongside front-row colleagues Nicky Smith and Dillon Lewis.

Tshiunza forges an all-Exeter lock partnership alongside Dafydd Jenkins, with fly-half Gareth Anscombe also back after injury.

Grady, meanwhile, is partnered in midfield by Johnny Williams, Leigh Halfpenny wins his 101st cap at full-back and another experienced campaigner – flanker Dan Lydiate – also features.